BIO 385 — Invertebrate Zoology
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Invertebrate Diversity
Phylum Arthropoda
Class Arachnida

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Arachnid Characteristics

  • Additional feeding appendages: pedipalps & maxilla
  • 4 pairs of walking legs
  • Lack abdominal appendages (except for spinnerets in some)
  • Nearly all are terrestrial
  • Respiration via book lungs or trachea

Class Arachnida


Subclass Acari

  • Ticks and mites
  • Most very tiny (< 1 mm), except for ticks
  • Some are external parasites
  • Very diverse; typically divided into two or more orders or superorders
    Parasitiformes are the ticks and various mostly parasitic mites
    Acariformes are a large, diverse group of mites (including parasitic forms)
  • May be polyphyletic
Hard Tick
Hard Tick, Dermacentor sp. (O. Parasitiformes); CA; a blood-feeding parasite
Dog Tick
American Dog Tick, Dermacentor variabilis (O. Parasitiformes); NE
Blue Oat Mite
Blue Oat Mite, Penthaleus sp. (O. Acariformes); AZ; feeds on plants
Hard Tick
Whirligig Mite, Anystis sp. (O. Acariformes); AZ; aquatic
Snout Mite
A predatory Snout Mite (Family Bdellidae, O. Acariformes), AZ
2 Mite species
Prostigmatid (left) & Oribatid (right) Mites (O. Acariformes); AZ; from leaf litter

Order Araneae

  • Spiders
  • 36,000 Species
  • Most are predators
  • Chelicerae modified into venom fangs
  • Silk glands and spinnerets
Black Widow Spider
Western Black Widow, Lactrodectus hesperus, female in ventral view
See also labeled photo.
Wolf Spider
Wolf Spider, Hogna sp. (Lycosidae); KY
See also labeled photo.
Tarantula
Chilean Rose Tarantula, Grammastola rosea; tarantulas are large, primitive spiders
Nursery Web Spider
Nursery Web Spider, Thaumasia sp.; Ecuador
Banded Agriope
Banded Garden Spider, Argiope trifasciata; Kenya
Crab Spider
Goldenrod Crab Spider, Misumena vatia; ME
Long-jawed Orb Weaver
Long-jawed Orb Weaver. Tetragnatha sp. (Tetragnathidae); AZ
Jumping Spider
Jumping Spider Phidippus sp. (Salticidae); AZ
Cellar Spider
Marbled Cellar Spider Holocnemus pluchei (Pholcidae); AZ (introduced from Mediterranean region)

Order Opiliones

  • Harvestmen & Daddy-Long-Legs
  • Prosoma & opisthosoma fused (no pedicel)
  • Small bodies but very long legs
  • Predators or scavengers
  • 1 Pair of eyes
  • Reduced chelicerae
Daddy Long Legs
Harvestman, Hadrobunus sp.?; KY
Desert Harvestman
Desert Harvestman, Eurybunus sp.; AZ
Harvestman
Ornate Harvestman, Vonones sayi; KY

Order Scorpiones

  • Scorpions
  • Abdomen divided into Mesosoma and "tail"
  • Large pedipalps form pincers (chelae)
  • Venomous stinger at end of tail
  • Reduced chelicerae
Northern Scorpion
Northern? Scorpion, Paruroctonus boreus?; UT
See also labeled photo.
Bark Scorpion ventral side
Arizona Bark Scorpion, Centruroides sculpturatus, ventral side of dried specimen
See also labeled photo.
AZ stripetail Scorpion
Arizona Striped-tail Scorpion, Vaejovis spinigerus; AZ
Sand Scorpion
Coachella Valley Sand Scorpion, Paruroctonus mesaensis; carrying young.
Uroctonus Scorpion
California Forest Scorpion, Uroctonus mordax; CA

Order Pseudoscorpiones

  • Pseudoscorpions
  • Large pedipalps form pincers (chelae)
  • Abdomen lacks elongated "tail" and stinger
  • Size tiny; often phoretic on beetles
Pseudoscorpion whole mount
Pseudoscorpion, whole mount microscope slide
Pseudoscorpion
Pseudoscorpion; AZ
Pseudoscorpion2
Pseudoscorpion; AZ

Order Solifugae or Solpugida

  • Sunspiders or Windscorpions
  • Very large forward-pointing chelicerae
  • Appear to have 5 pairs of legs: the anterior pair is actually enlarged leg-like pedipalps used as feelers
  • Fast-running predators
Wind Scorpion
Straight-faced Wind Scorpion, Eremobates sp.?, male; AZ
Wind Scorpion female
Straight-faced Wind Scorpion, Eremobates sp.?, female; AZ
Sun Spider
Sun Spider, Eremocosta sp.?; dried specimen

Order Amblypygi

  • Tailless Whipscorpions
  • Pedipalps with spikes but not jointed chelae
  • Flattened body
  • First pair of walking legs extremely long (at least several times length of other legs) and function as feelers
  • Lack telson
Sonoran whipscorpion
Sonoran Whipscorpion, Paraphrynus mexicanus (native to southern Arizona)
Whipscorpion
Florida Whipscorpion, Phrynus marginemaculatus; FL
Whip Spider
African Whipspider, Damon variegatus

Order Uropygi

  • Whipscorpions or Vinegaroons
  • Enlarged pedipalps but with reduced chelae
  • Flattened body in most
  • First pair of walking legs longer and slenderer than others, act as feelers
  • Posterior end with a thin, straight telson that squirts acetic acid
  • Short-tailed whipscorpions are small (<5 mm), tropical species with very short telson; sometimes placed in their own order, Schizomyida
Vinegaroon
Philippine Vinegaroon, Minbosius manilanus
Vinegaroon
Giant Vinegaroon, Mastigoproctus giganteus
Shorttailed Whipscorpion
Short-tailed Whipscorpion, Schizomus floridanus, preserved specimen
This page last updated 6 June 2021 by Udo M. Savalli ()
Images and text © Udo M. Savalli. All rights reserved.